Trogoderma

Trogoderma
	Khapra beetle (Trogoderma granarium)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Dermestidae
Tribe:	Megatomini
Genus:	Trogoderma; Dejean, 1821

Trogoderma is a genus of beetles in the family Dermestidae, the skin beetles. There are about 135 species worldwide.^[1]

Some species are pests of stored animal and plant products.^[2]

Taxonomy

Recent research shows that Trogoderma (in its broad, traditional definition) is polyphyletic. Many species in the Southern Hemisphere are genetically distinct from the Holarctic species, which led to their reclassification under the genus Eurhopalus (Zhou et al., 2022).^[3] However, this change is not widely adopted, which in some cases is caused by differing opinions on taxonomy - e.g., the World Dermestidae Catalogue (Jiří Háva, 2023) still recognizes the old classification system and actively reinstated several genera synonymized with Eurhopalus.^[4]

Species

Trogoderma adelaideum Blackburn, 1891
Trogoderma albonotatum Reiche, 1868
Trogoderma alpicorum Blackburn, 1891
Trogoderma americanum Dejean, 1837
Trogoderma angustum Solier, 1849
Trogoderma antennale Broun, 1893
Trogoderma antipodum Blackburn, 1891
Trogoderma apicale MacLeay, 1871
Trogoderma apicipenne Reitter, 1881
Trogoderma arcanum Zhantiev, 2002
Trogoderma armstrongi Háva, 2003
Trogoderma asperatum Fauvel, 1903
Trogoderma atrum Philippi & Philippi, 1864
Trogoderma attagenoides Pascoe, 1866
Trogoderma bactrianum Zhantiev, 1970
Trogoderma baeri Pic, 1915
Trogoderma ballfinchae Beal, 1954
Trogoderma beali Mroczkowski, 1968
Trogoderma bicinctum Reitter, 1881
Trogoderma blackburni Lea, 1908
Trogoderma boganense Armstrong, 1942
Trogoderma caboverdianum Kalík, 1986
Trogoderma callubriense Armstrong, 1945
Trogoderma carteri Armstrong, 1942
Trogoderma cavum Beal, 1982
Trogoderma celatum Sharp, 1902
Trogoderma consors Arrow, 1915
Trogoderma davidsoni Háva & Kadej, 2006
Trogoderma debilius Blackburn, 1903
Trogoderma denticorne Normand, 1936
Trogoderma deserti Beal, 1954
Trogoderma difficile Blackburn, 1891
Trogoderma dominicanum Háva & Kadej, 2006
Trogoderma ellipticum Armstrong, 1942
Trogoderma explanaticolle Armstrong, 1942
Trogoderma exsul Blackburn, 1903
Trogoderma eyrense Blackburn, 1891
Trogoderma fantasticum Háva & Kadej, 2006
Trogoderma fasciferum Blatchley, 1914
Trogoderma frater Arrow, 1915
Trogoderma froggatti Blackburn, 1892
Trogoderma glabrum (Herbst, 1783)
Trogoderma granarium Everts, 1898 – Khapra beetle
Trogoderma granulatum Broun, 1886
Trogoderma grassmani Beal, 1954
Trogoderma halsteadi Veer & Rao, 1994
Trogoderma hobartense Armstrong, 1942
Trogoderma impressiceps Pic, 1915
Trogoderma inclusum LeConte, 1854
Trogoderma inconspicuum Armstrong, 1942
Trogoderma insulare Chevrolat, 1863
Trogoderma irroratum Reitter, 1881
Trogoderma kaliki Háva, 2006
Trogoderma koenigi Pic, 1954
Trogoderma laevipenne Armstrong, 1942
Trogoderma larvale Háva, Prokop & Herrmann, 2006
Trogoderma leai Armstrong, 1942
Trogoderma lindense Blackburn, 1891
Trogoderma longisetosum Chao & Lee, 1966
Trogoderma longius Blackburn, 1903
Trogoderma macleayi Blackburn, 1891
Trogoderma madecassum Pic, 1924
Trogoderma maderae Beal, 1954
Trogoderma maestum Broun, 1880
Trogoderma marginicolle Armstrong, 1942
Trogoderma mauricepici Háva, 2003
Trogoderma maurulum Blackburn, 1903
Trogoderma megatomoides Reitter, 1881
Trogoderma melanarium Sturm, 1843
Trogoderma mexicanum Reitter, 1881
Trogoderma meyricki Blackburn, 1891
Trogoderma mongolicum Zhantiev, 1973
Trogoderma morio Erichson, 1842
Trogoderma nigrobrunneum Armstrong, 1942
Trogoderma nigronitidum Armstrong, 1945
Trogoderma nitens Arrow, 1915
Trogoderma obscurum Pic, 1936
Trogoderma occidentale Blackburn, 1891
Trogoderma octaedron Peyerimhoff, 1943
Trogoderma okumurai Beal, 1964
Trogoderma ornatum (Say, 1825)
Trogoderma paralia Beal, 1954
Trogoderma parvum Armstrong, 1942
Trogoderma pectinicornis Reitter, 1881
Trogoderma peruvianum Pic, 1954
Trogoderma picinum Armstrong, 1945
Trogoderma plagifer Casey, 1916
Trogoderma primum (Jayne, 1882)
Trogoderma punctatum Broun, 1886
Trogoderma puncticolle Broun, 1914
Trogoderma quadrifasciatum Broun, 1893
Trogoderma reitteri Blackburn, 1892
Trogoderma rubiginosum Solier, 1849
Trogoderma ruficolle Reitter, 1881
Trogoderma rufipenne Armstrong, 1942
Trogoderma rufonotatum Pic, 1942
Trogoderma rufopictum Arrow, 1915
Trogoderma schawalleri Háva, 2007
Trogoderma schmorli Reitter, 1881
Trogoderma seminigrum Pic, 1915
Trogoderma serraticorne (Fabricius, 1792)
Trogoderma serratum Dejean, 1837
Trogoderma serrigerum Sharp, 1877
Trogoderma setulosum Armstrong, 1942
Trogoderma signatum Sharp, 1877
Trogoderma silvicolum Armstrong, 1949
Trogoderma simplex Jayne, 1882
Trogoderma sinense Pic, 1927
Trogoderma singulare Blackburn, 1891
Trogoderma sinistrum Fall, 1926
Trogoderma socium Lea, 1895
Trogoderma stachi Mroczkowski, 1958
Trogoderma sternale Jayne, 1882
Trogoderma subfascia (Dahl, 1823)
Trogoderma subrotundatum Reitter, 1881
Trogoderma subtile Reitter, 1881
Trogoderma tasmanicum Armstrong, 1942
Trogoderma tenuefasciatum Reitter, 1881
Trogoderma teukton Beal, 1956
Trogoderma thoracicum Reitter, 1881
Trogoderma tolarnense Blackburn, 1903
Trogoderma unifasciatum Pic, 1942
Trogoderma variabile Ballion, 1878
Trogoderma variegatum (Solier, 1849)
Trogoderma varipes Blackburn, 1892
Trogoderma varium Matsumura & Yokoyama, 1928
Trogoderma versicolor (Creutzer, 1799)
Trogoderma vicinum Dejean, 1837
Trogoderma villosum Dejean, 1821
Trogoderma vulneratum Fauvel, 1903
Trogoderma westerduijni Háva & Herrmann, 2007
Trogoderma whitei Armstrong, 1942
Trogoderma yorkense Blackburn, 1891
Trogoderma yunnaeunse Zhang & Liu, 1986

References

^ Háva, J. (2009). Review of the genus Trogoderma from Madagascar (Coleoptera: Dermestidae: Megatominae). Baltic Journal of Coleopterology, 9(2), 111-117.
^ Wei, C., et al. (2015). Scanning electron microscope observations on the antennal sensilla of two stored grain pests Trogoderma granarium and Trogoderma variabile (Coleoptera: Dermestidae). Florida Entomologist, 98(1), 140-148.
^ Yu-Lingzi Zhou; James A Nicholls; Zhen-Hua Liu; Diana Hartley; Andras Szito; Adam Ślipiński; Andreas Zwick (1 November 2022). "Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma granarium Everts" (PDF). Insect Systematics and Diversity. 6 (6). doi:10.1093/ISD/IXAC026. ISSN 2399-3421. Wikidata Q116673277. Archived from the original (PDF) on 2 July 2024.
^ Háva, Jiří (2023). "World Dermestidae. Subfamily Megatominae" (PDF). Archived from the original (PDF) on 2 June 2024. Retrieved 12 June 2024.

[Trogoderma_madagascar-1] Háva, J. (2009). Review of the genus Trogoderma from Madagascar (Coleoptera: Dermestidae: Megatominae). Baltic Journal of Coleopterology, 9(2), 111-117.

[wei-2] Wei, C., et al. (2015). Scanning electron microscope observations on the antennal sensilla of two stored grain pests Trogoderma granarium and Trogoderma variabile (Coleoptera: Dermestidae). Florida Entomologist, 98(1), 140-148.

[Zhou_et_al_2022-3] Yu-Lingzi Zhou; James A Nicholls; Zhen-Hua Liu; Diana Hartley; Andras Szito; Adam Ślipiński; Andreas Zwick (1 November 2022). "Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma granarium Everts" (PDF). Insect Systematics and Diversity. 6 (6). doi:10.1093/ISD/IXAC026. ISSN 2399-3421. Wikidata Q116673277. Archived from the original (PDF) on 2 July 2024.

[wd-species-4] Háva, Jiří (2023). "World Dermestidae. Subfamily Megatominae" (PDF). Archived from the original (PDF) on 2 June 2024. Retrieved 12 June 2024.

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