Sea snail

Determining whether some gastropods should be called sea snails is not always easy. Some species that live in brackish water (such as certain neritids) can be listed as either freshwater snails or marine snails, and some species that live at or just above the high tide level (for example, species in the genus Truncatella) are sometimes considered to be sea snails and sometimes listed as land snails.

Sea snails are a very large and diverse group of animals. Most snails that live in salt water respire using a gill or gills; a few species, though, have a lung, are intertidal, and are active only at low tide when they can move around in the air. These air-breathing species include false limpets in the family Siphonariidae and another group of false limpets in the family Trimusculidae.

Many, but not all, sea snails have an operculum.

The shells of most species of sea snails are spirally coiled. Some, though, have conical shells, and these are often referred to by the common name of limpets. In one unusual family (Juliidae), the shell of the snail has become two hinged plates closely resembling those of a bivalve; this family is sometimes called the "bivalved gastropods".

Their shells are found in a variety of shapes and sizes, but are normally very small. Those living species of sea snails range in size from Syrinx aruanus, the largest living shelled gastropod species at 91 cm (36 in), to minute species whose shells are less than 1 mm at adult size. Because the shells of sea snails are strong and durable in many cases, as a group they are well represented in the fossil record.

The shells of snails are complex and grow at different speeds. The speed of growth is affected by a few variables such as the temperature of the water, depth of the water, food present for the snail, as well as isotopic oxygen levels. By looking at the composition of aragonite in the growth layers of mollusks you can predict the size the mollusk shell can reach.^[1]

The following cladogram is an overview of the main clades of living gastropods based on the taxonomy of Bouchet & Rocroi (2005),^[2] with taxa that contain saltwater or brackish water species marked in boldface (some of the highlighted taxa consist entirely of marine species, but some of them also contain freshwater or land species.)

• Clade Patellogastropoda
• Clade Vetigastropoda
• Clade Cocculiniformia
• Clade Neritimorpha
  • Clade Cycloneritimorpha
• Clade Caenogastropoda
  • Informal group Architaenioglossa
  • Clade Sorbeoconcha
  • Clade Hypsogastropoda
    • Clade Littorinimorpha
    • Informal group Ptenoglossa
    • Clade Neogastropoda
• Clade Heterobranchia
  • Informal group Lower Heterobranchia
  • Informal group Opisthobranchia
    • Clade Cephalaspidea
    • Clade Thecosomata
    • Clade Gymnosomata
    • Clade Aplysiomorpha
    • Group Acochlidiacea
    • Clade Sacoglossa
    • Group Cylindrobullida
    • Clade Umbraculida
    • Clade Nudipleura
      • Clade Pleurobranchomorpha
      • Clade Nudibranchia
        • Clade Euctenidiacea
        • Clade Dexiarchia
          • Clade Pseudoeuctenidiacea
          • Clade Cladobranchia
            • Clade Euarminida
            • Clade Dendronotida
            • Clade Aeolidida
  • Informal group Pulmonata
    • Informal group Basommatophora
    • Clade Eupulmonata
      • Clade Systellommatophora
      • Clade Stylommatophora
        • Clade Elasmognatha
        • Clade Orthurethra
        • Informal group Sigmurethra

A number of species of sea snails are harvested in aquaculture and used by humans for food, including abalone, conch, limpets, whelks (such as the North American Busycon species and the North Atlantic Buccinum undatum) and periwinkles including Littorina littorea.

The shells of sea snails are often found washed up on beaches. Because many are attractive and durable, they have been used to make necklaces and other jewelry since prehistoric times.

The shells of a few species of large sea snails within the Vetigastropoda have a thick layer of nacre and have been used as a source of mother of pearl. Historically, the button industry relied on these species for a number of years.

The shells of sea snails are used for protection by many kinds of hermit crabs. A hermit crab carries the shell by grasping the central columella of the shell using claspers on the tip of its abdomen.

• Freshwater snail
• Terrestrial molluscs
• Land snail
• Sea slug