Chimpanzee

Pan troglodytes[1] Temporal range: 4–0 mya[2]
P. t. schweinfurthii in Kibale National Park, Uganda
Conservation status
Endangered  (IUCN 3.1)[3]
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Primates
Suborder:	Haplorhini
Infraorder:	Simiiformes
Family:	Hominidae
Subfamily:	Homininae
Tribe:	Hominini
Genus:	Pan
Species:	P. troglodytes
Binomial name
Pan troglodytes (Blumenbach, 1775)
Subspecies
Pan troglodytes verus Pan troglodytes ellioti Pan troglodytes troglodytes Pan troglodytes schweinfurthii
Distribution of subspecies Pan troglodytes verus (green) P. t. ellioti (grey) P. t. troglodytes (red) P. t. schweinfurthii (blue)
Synonyms
Simia troglodytes Blumenbach, 1775 Troglodytes troglodytes (Blumenbach, 1776) Troglodytes niger E. Geoffroy, 1812 Pan niger (E. Geoffroy, 1812) Anthropopithecus troglodytes (Sutton, 1883)

The chimpanzee (Pan troglodytes), or robust chimpanzee, is a species of great ape. The common chimpanzee is often called the chimpanzee (or "chimp"), but this can refer to both species in the genus Pan: the common chimpanzee and the closely related bonobo.

Evidence from fossils and DNA sequencing show both species of chimpanzees are the closest living relatives to modern humans.

The common chimpanzee is more robust than the bonobo, weighing between 40 and 65 kg (88 and 143 lb) and measuring approximately 1.3 to 1.6 m (4 ft 3 in to 5 ft 3 in). Its gestation period is eight months. The infant is weaned at about three years old, but usually maintains a close relationship with its mother for several more years; it reaches puberty at the age of eight to 10, and its lifespan in captivity is about 50 years.

The common chimpanzee lives in groups which range from 15 to 150 members, although individuals travel and forage in much smaller groups during the day. The species lives in a male-dominated strict hierarchy. Disputes can generally be settled without the need for violence.

Nearly all chimpanzee populations have been seen using tools, modifying sticks, rocks, grass, and leaves. They use them for getting honey, termites, ants, nuts, and water. The species has also been seen making sharpened sticks to spear Senegal bushbabies out of small holes in trees. So although it is mainly a herbivore, it does eat other animals to some extent.

The common chimpanzee is listed on the IUCN Red List as an endangered species. There are between 170,000 and 300,000 individuals. They live in the forests and savannahs of West and Central Africa. The biggest threats to the common chimpanzee are habitat destruction, poaching and disease.

Jane Goodall discovered that chimps hunt and eat smaller primates such as Colobus monkeys. Goodall watched a hunting group isolate a monkey high in a tree, block all possible exits. Then one chimpanzee climbed up and captured and killed the monkey.^[4] The others then each took parts of the carcass, sharing with other members of the troop in response to begging behaviours.^[4] The chimps at Gombe kill and eat as much as one-third of the Colobus population in the park each year.^[5] This was a major scientific discovery: it challenged previous ideas of chimp diet and behavior.

"Goodall’s Gombe data have also led researchers to take a closer look at the role that hunting plays in chimp feeding habits. One recent Gombe study, for instance, concluded that the 45 members of one troop ate a ton of monkey meat per year. During one hunting binge, chimps killed 71 colobus monkeys in 68 days; one chimp alone killed 42 monkeys over five years. All told, chimps may kill and eat a third of the Gombe’s Colobus population each year. Researchers have also found that lower-ranking males often trade the meat for mating privileges; such trades may help prevent inbreeding by keeping a single group of males from fathering the majority of a troop’s children".^[5]

Also startling, and disturbing, was the tendency for aggression and violence within chimpanzee troops. Goodall observed dominant females deliberately killing the young of other females in the troop in order to maintain their dominance,^[5] sometimes going as far as cannibalism.^[6][7]

She says of this revelation, "During the first ten years of the study I had believed […] that the Gombe chimpanzees were, for the most part, rather nicer than human beings. […] Then suddenly we found that chimpanzees could be brutal—that they, like us, had a darker side to their nature".^[6] These findings revolutionized our knowledge of chimpanzee behaviour. They were further evidence of the social similarities between humans and chimpanzees, albeit in a much darker manner.

If they can, male chimpanzees try to kill the male members of neighbouring groups. Males work together when they spot a chance to make a lightning raid on an isolated male from the other group. They kill him. In Gombe, Tanzania, a group in the 1970s was seen to kill seven of their neighbours one by one, until all were gone. It can take years for this to happen but, when it does, the remaining females and the neighbouring territory are added to the now larger group. Attacks like this are carefully planned, done only when success is likely, and carried out in silence. Their behaviour is quite different from any other behaviour known in chimpanzees:

"About every two weeks, males are drawn by some unknown signal to walk very quietly, single file, into a neighbouring territory to attack a vulnerable male... the overwhelming targets are other males".^[8]249[9]

The advantage for the males that win is to breed more children. Their tribe also holds a larger territory, and so has access to more food.^[10] Several authors have drawn a connection between this behaviour and the origins of human warfare.^{[8][11][12][13]}

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