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{{Automatic taxobox
| name = ''Pan''<ref name=MSW3>{{MSW3 Groves|pages=182–3|id=12100796}}</ref>
| fossil_range = Middle [[Pliocene]] – [[Holocene|present]]
| image = Composite image of male chimpanzee (left) and male bonobo (right).jpg
| image_caption = Members of the genus ''Pan'': chimpanzee (left) and bonobo (right)
| taxon = Pan
| authority = [[Lorenz Oken|Oken]], 1816
| type_species = ''[[Common chimpanzee|Simia troglodytes]]'' = ''Pan troglodytes''
| type_species_authority = [[Johann Friedrich Blumenbach]], 1776
| subdivision_ranks = [[Species]]
| subdivision = ''[[Common chimpanzee|Pan troglodytes]]''<br/>
''[[Bonobo|Pan paniscus]]''
| range_map = Pan (genus) distribution map.png
| range_map_caption = Distribution of ''Pan troglodytes'' (common chimpanzee) and ''Pan paniscus'' (bonobo, in red)
| synonyms = ''Troglodytes'' <small>[[Étienne Geoffroy Saint-Hilaire|E. Geoffroy]], 1812 ([[Troglodytes (bird)|preoccupied]])</small><br/>
''Mimetes'' <small>Leach, 1820 ([[Noctua (moth)|preoccupied]])</small><br/>
''Theranthropus'' <small>Brookes, 1828</small><br/>
''Chimpansee'' <small>Voight, 1831</small><br/>
''[[Anthropopithecus]]'' <small>[[Henri Marie Ducrotay de Blainville|Blainville]], 1839</small><ref>Bernard Wood ''et alii'', ''Wiley-Blackwell Encyclopedia of Human Evolution'', June 2013 (single-volume paperback version of the original 2011 2-volume edition), 1056 pp.; {{ISBN|978-1-1186-5099-8}}</ref><br/>
''Hylanthropus'' <small>Gloger, 1841</small><br/>
''Pseudanthropus'' <small>Reichenbach, 1862</small><br/>
''Engeco'' <small>Haeckel, 1866</small><br/>
''Fsihego'' <small>DePauw, 1905</small>
}}
The genus '''''Pan''''' is made up of two living species: the [[chimpanzee]] and the [[bonobo]]. Taxonomically, these two ape species are called '''''panins''''';<ref>{{cite book |last= Muehlenbein|first= M. P.|date= 2015|title= Basics in Human Evolution|url= https://www.elsevier.com/books/basics-in-human-evolution/muehlenbein/978-0-12-802652-6 |publisher= Elsevier Science |pages= 114–115|isbn= 9780128026526}}</ref><ref>{{cite news|title=Bonobo anatomy reveals stasis and mosaicism in chimpanzee evolution, and supports bonobos as the most appropriate extant model for the common ancestor of chimpanzees and humans|url= https://www.nature.com/articles/s41598-017-00548-3|access-date=January 10, 2019|publisher=nature.com|date=April 4, 2017}}</ref> however, both species are more commonly called '''chimpanzees''' or '''chimps'''. Together with [[gorilla]]s, [[orangutan]]s and [[human]]s, they are part of the family [[Hominidae]].

==Taxonomy==
* Genus ''Pan'': Chimpanzees
** [[Common chimpanzee]], ''[[Pan troglodytes]]''
** Bonobo or Pygmy chimpanzee, ''Pan paniscus''

==Habitat==
The common chimpanzee lives in West and Central [[Africa]]. The bonobo lives in the [[rain forest]]s of the [[Democratic Republic of the Congo]]. The two species are on opposite sides of the [[Congo River]].

==Life and description==
Chimpanzees mainly eat fruit, leaves, flowers, seeds, bark, honey, insects, bird eggs, and meat. They spend a lot of time with other chimpanzees from their group, acting up, playing, and chatting. Sometimes they will groom each other; combing and looking through each other's thick fur; picking out the dirt and insects. [[Groom]]ing helps chimps feel comfortable and friendly.

Chimpanzees can walk on two feet, but they prefer to move about on all four legs. On the ground, they [[knuckle-walk|walk on their hind feet and knuckles]].<ref name="Nature">{{cite news|title=These fists were made for walking|publisher=Nature|date=March 23, 2000|url=https://www.nature.com/articles/news000323-7|access-date=July 8, 2020|author=Henry Gee}}</ref><ref name="Ringer">{{cite news|url=https://thedaily.case.edu/arent-humans-knuckle-walkers/|publisher=The Daily|access-date=July 7, 2020|title=Why aren't humans 'knuckle-walkers'?|date=April 9, 2018}}</ref><ref name="Conv">{{cite journal|title=Knuckle-walking Anteater: A Convergence Test of Adaptation for Purported Knuckle-Walking Features of African Hominidae|journal=Am J Phys Anthropol|year=2005|volume=128|issue=3|pages=639–58|doi=10.1002/ajpa.20192|access-date=July 8, 2020|author=Caley M Orr|pmid=15861420|url=https://pubmed.ncbi.nlm.nih.gov/15861420/}}</ref> They have hands that look like human hands, but their thumbs are shorter than those of humans. At night, chimpanzees sleep in nests that they make on tree branches. They bend twigs and tuck in leaves to make a soft platform to rest in a place that is safe from [[enemies]] on the ground. The [[gestation]] period of chimpanzees lasts between six and eight months. Usually only one offspring is produced; they rarely have twins. Chimpanzees live up to 60 years in the wild.<ref name=chimp>{{cite book |author= Pous, Dinorah|title= Blue Planet English through Science level 5|url= https://archive.org/details/blueplanetlevel50000unse|year=2003|publisher= Alpha Omega Publications|isbn= 970-10-3769-3}}</ref>

[[Jane Goodall]] has studied chimpanzees since 1960. She observed how chimpanzees use tools in several ways. They will pick up rocks to crack nuts for a meal. They also strip down twigs and stick them into a [[termite]] mound to collect a tasty snack. Some have been known to make a sponge from leaves in order to hold more drinking water. The chimp [[wikt:chew|chews]] leaves to soften them up, dips them in rainwater, and then squeezes the water into its mouth. Chimpanzees are also known to think ahead and solve problems.<ref>Köhler, Wolfgang 1925. ''The mentality of apes'', transl. from the 2nd German edition by Ella Winter. Kegan, Trench: London and Harcourt, Brace and World: New York. Original was ''Intelligenzprüfungen an Anthropoiden'', Berlin 1917.</ref><ref name="Goodall">The Jane Goodall Institute: [http://www.janegoodall.org/chimp_central/default.asp "Chimpanzee Central"], 2008.</ref>

==Chimpanzee aggression==
The behavior described in this section refers to the common chimpanzee. At present there is no evidence that the bonobo has a similar level of aggressive behavior.

===Attacking monkeys===
Chimpanzees attack ''[[Colobus]]'' monkeys by working as a team to corner them in the high branches of the trees. Then they tear the monkey apart and eat it.<ref name=PBS>{{cite web |publisher=[[PBS]] |url=https://www.pbs.org/wnet/nature/episodes/jane-goodalls-wild-chimpanzees/introduction/1908/ |title=Jane Goodall's Wild Chimpanzees |year=1996 |access-date=2010-07-28}}</ref> It is thought the main benefit is that meat is a more rich source of [[nutrition]] than their usual vegetarian diet.
:"Goodall’s Gombe data have also led researchers to take a closer look at the role that hunting plays in chimp feeding habits. One recent Gombe study, for instance, concluded that the 45 members of one troop ate a ton of monkey meat per year. During one hunting binge, chimps killed 71 colobus monkeys in 68 days; one chimp alone killed 42 monkeys over five years. All told, chimps may kill and eat a third of the Gombe’s colobus population each year. Researchers have also found that lower-ranking males often trade the meat for mating privileges; such trades may help prevent inbreeding".<ref name=PBS /> Otherwise one or two males might father most of a troop’s children.

===Attacking other chimpanzee groups===
If they can, male chimpanzees try to kill the male members of neighboring groups. Males work together when they spot a chance to make a lightning raid on an isolated male from the other group. They kill him. In [[Gombe Stream National Park|Gombe]], [[Tanzania]], a group in the 1970s was seen to kill seven of their neighbors one by one, until all were gone. It can take years for this to happen but, when it does, the remaining females and the neighboring territory are added to the now larger group. Attacks like this are carefully planned, done only when success is likely, and carried out in silence.

The advantage for the males that triumph is to breed more children. Their tribe also holds a larger territory, and so has access to more food.<ref>Mitani J.C; Watts D.P. & Amsler S.J. 2010. Lethal intergroup aggression leads to territorial expansion in wild chimpanzees. ''Current Biology'' '''20''' (12) R507/8.</ref> Several authors have drawn a connection between this behaviour and the origins of human warfare.<ref>Wrangham R. & Peterson D. 1996. Demonic males: apes and the origins of human warfare. Boston: Houghton Mifflin.</ref><ref>Wrangham R. 2006. Why apes and humans kill. In Jones M. & Fabian A. (eds) ''Conflict''. Cambridge University Press.</ref><ref>Trivers, Robert 2011. ''Deceit and self-deception: fooling yourself the better to fool others''. London: Penguin, p249/250: Chimpanzee raiding > human warfare. {{ISBN|978-0-141-01991-8}}</ref>

Scientists from the Max Planck Institute for Evolutionary Anthropology studied chimpanzees in Taï National Park in [[Côte d’Ivoire]]. The project followed four groups of chimpanzees for twenty years. They found that female chimpanzees patrolled territory and fought with chimpanzees from other groups too, not just males. The scientists wondered if human beings have been imagining that male chimpanzees do all the fighting because we think of human warfare being done mostly by men.<ref name="Female">{{cite press release|title=In chimpanzees, females contribute to the protection of the territory|date=May 27, 2020|url=https://www.mpg.de/14884914/0527-evan-019609-female-chimpanzees-are-also-important-in-helping-to-win-and-keep-a-territory|author=Max Planck Institute for Evolutionary Anthropology|access-date=May 28, 2020}}</ref><ref name="Eureka">{{cite press release|title=In chimpanzees, females contribute to the protection of the territory|url=https://www.eurekalert.org/pub_releases/2020-05/mpif-icf052720.php|publisher=Eurekalert|date=May 27, 2020|access-date=May 28, 2020|author=Max Planck Institute for Evolutionary Anthropology|archive-date=June 6, 2020|archive-url=https://web.archive.org/web/20200606034423/https://www.eurekalert.org/pub_releases/2020-05/mpif-icf052720.php|url-status=dead}}</ref><ref name="FemalesContribute">{{cite journal|doi=10.1098/rsos.200577 |author1=Sylvia Lemoine|author2=Christophe Boesch|author3= Anna Preis|author4= Liran Samuni|author5= Catherine Crockford|author6=Roman M. Wittig|title=Group dominance increases territory size and reduces neighbour pressure in wild chimpanzees|journal=Royal Society Open Science|date=May 27, 2020|volume=7|issue=5|page=200577|pmid=32537232|pmc=7277268|bibcode=2020RSOS....700577L}}</ref>
:One of the scientists, Catherine Crockford, said: "We are now accumulating compelling evidence that cooperation among non-related individuals, and among males and females, has likely been under selection due to the competition between neighboring groups. These findings have therefore strong implications to understand the evolution of cooperation in social species and in humans in particular. Continuous and long-lasting research and conservation efforts are necessary if we want to understand the interplay between competition and cooperation in this emblematic species for human evolution, and how it relates to our own evolution."<ref name="Female" />

==Communication==
Chimpanzees show their emotions with their faces and sounds. They make hooting sounds to [[zoosemiotics|express]] the discovery of food, and the face of a chimpanzee with a scowling face and lips pressed is to express annoyance. This means the chimpanzee may attack. Or, the chimpanzee may bare its teeth to express that it is afraid or that a more [[dominant]] chimp is approaching.<ref name=chimp /><sup>p74/5</sup>

==Relationship with humans==
According to a [[genome]] study done by the Chimpanzee Sequencing and Analysis Consortium, humans share either 96% or 95% of their [[DNA]] with chimpanzees. However, this applies only to [[single nucleotide polymorphism]]s, that is, changes in ''single'' base pairs only. The full picture is rather different.

24% of the chimpanzee genome does not align with the human genome, and so cannot be directly compared. There are 3% further alignment gaps, 1.23% SNP differences, and 2.7% copy number variations totaling at least 30% differences between chimpanzee and ''Homo sapiens'' genomes. On the other hand, 30% of all human proteins are identical in sequence to the corresponding chimpanzee proteins.<ref name=Consortium>{{cite journal | author = The Chimpanzee Sequencing and Analysis Consortium | year = 2005 | title = Initial sequence of the chimpanzee genome and comparison with the human genome | journal = Nature | volume = 437 | issue = 1 September 2005 | pages = 69–87 | doi = 10.1038/nature04072 | pmid = 16136131|bibcode = 2005Natur.437...69.| s2cid = 2638825}}</ref>

==Related pages==
* [[Ape]]

==References==
{{reflist}}

{{wikispecies|Pan}}
{{Commons category|Pan (genus)|Pan}}

[[Category:Hominins]]

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