Abstract
Turanoceratops tardabilis from the Upper Cretaceous (Turonian) Bissekty Formation of Dzharakuduk, Uzbekistan, represents the first definite ceratopsid dinosaur recorded from Asia. Reexamination of the original and study of newly collected material indicate that Turanoceratops has well-developed supraorbital horns and apparently lacks a nasal horn. Turanoceratops is more derived than the more or less coeval Zuniceratops from the Moreno Hill Formation of New Mexico in the presence of double-rooted teeth and of two or three teeth in each vertical dental file.




References
Brown B (1917) A complete skeleton of the horned dinosaur Monoclonius, and description of a second skeleton showing skin impressions. Bull Am Mus Nat Hist 37:281–306
Chinnery BJ, Weishampel DB (1998) Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) and relationships among basal neoceratopsians. J Vert Paleontol 18:569–585
Coombs WP Jr (1979) Osteology and myology of the hindlimb in the Ankylosauria. J Paleontol 53:666–684
Dodson P, Forster CA, Sampson SD (2004) Ceratopsidae. In: Weishampel DB, Dodson P, Osmólska H (eds) The Dinosauria, 2nd edn. University of California Press, Berkeley, pp 494–513
Forster CA (1996) New information on the skull of Triceratops. J Vert Paleontol 16:246–258
Goloboff P (1999) NONA (No Name) version 2. Published by the author, Tucumán, Argentina. Available online at www.cladistics.org
Hatcher JB, Marsh OC, Lull RS (1907) The Ceratopsia. US Geol Surv Monogr 49:1–300
Lehman TM (1989) Chasmosaurus mariscalensis, sp. nov., a new ceratopsian dinosaur from Texas. J Vert Paleontol 9:137–162
Lull RS (1933) A revision of the Ceratopsia or horned dinosaurs. Mem Peabody Mus Nat Hist Yale Univ 3(3):1–175
Makovicky PJ, Norell MA (2006) Yamaceratops dorngobiensis, a new primitive ceratopsian (Dinosauria, Ornithischia) from the Cretaceous of Mongolia. Am Mus Novit 3530:1–42
Marsh OC (1888) A new family of horned dinosaurs from the Cretaceous. Am J Sci (3) 36:477–478
Marsh OC (1890) Additional characters of the Ceratopsidae, with notice of new Cretaceous dinosaurs. Am J Sci (3) 39:418–426
Nessov LA (1986) First discovery of the Late Cretaceous bird Ichthyornis in the Old World and some other bones of birds from the Cretaceous and Paleogene of Middle Asia. Trudy Zool Inst AN SSSR 147:31–38 In Russian
Nessov LA (1988) Assemblages of vertebrates of the late Mesozoic and Paleocene of Middle Asia. Trudy XXXI Sess Vsesoyuz Paleont Obshchestva. Nauka, Leningrad, pp 93–101 In Russian
Nessov LA (1995) Dinosaurs of northern Eurasia: new data about assemblages, ecology and paleobiogeography. Izdatelstvo Sankt-Peterburgskogo Universiteta, St. Petersburg In Russian
Nessov LA (1997) Cretaceous nonmarine vertebrates of northern Eurasia. Izdatel’stvo Sankt-Peterburgskogo Universiteta, Saint Petersburg In Russian
Nessov LA, Kaznyshkina LF, Cherepanov GO (1989) Ceratopsian dinosaurs and crocodiles of the Mesozoic of Middle Asia. In: Bogdanova TN, Khozatsky LI (eds) Theoretical and applied aspects of modern paleontology. Nauka, Leningrad, pp 144–154 In Russian
Nixon KC (1999) WinClada version 1.0000. Published by the author, Ithaca, NY. Available online at www.cladistics.org
Owen R (1842) Report on British fossil reptiles. Part II. Rep Brit Ass Adv Sci 1841:60–204
Penkalski P, Dodson P (1999) The morphology and systematics of Avaceratops, a primitive horned dinosaur from the Judith River Formation (late Campanian) of Montana, with the description of a second skull. J Vert Paleontol 19:692–711
Riabinin AN (1931) On dinosaurian remains from the Upper Cretaceous of the lower parts of Amu-Daria River. Zap Russ Min Obshch (2) 60:114–118 In Russian
Ryan M, Currie PJ (1998) First report of protoceratopsians (Neoceratopsia) from the Late Cretaceous Judith River Group, Alberta, Canada. Can J Earth Sci 35:820–826
Seeley HG (1887) On the classification of the fossil animals commonly named Dinosauria. Proc R Soc Lond 43:165–171
Sereno PC (1997) The origin and evolution of dinosaurs. Ann Rev Earth Planet Sci 25:435–489
Sereno PC (2000) The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia. In: Benton MJ, Shishkin MA, Unwin DM, Kurochkin EN (eds) The age of dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge, pp 480–516
Wolfe DG, Kirkland JI (1998) Zuniceratops christopheri n. gen., n. sp., a ceratopsian dinosaur from the Moreno Hill Formation (Cretaceous, Turonian) of west-central New Mexico. New Mexico Mus Nat Hist Sci Bull 14:303–317
You HL, Dodson P (2003) Redescription of neoceratopsian dinosaur Archaeoceratops and early evolution of Neoceratopsia. Acta Palaeont Pol 48:261–272
You HL, Dodson P (2004) Basal Ceratopsia. In: Weishampel DB, Dodson P, Osmólska H (eds) The Dinosauria, 2nd edn. University of California Press, Berkeley, pp 478–493
Acknowledgments
Fieldwork for this study was funded by grants from the National Science Foundation (EAR-9804771 and 0207004) and National Geographic Society (5901-97 and 6281-98) to J. D. Archibald and H.-D.S. and US Civilian Research and Development Foundation grant RUB1-2860-ST-07 to A.A. and J. D. Archibald. We thank the National Academy of Sciences of Uzbekistan, especially D. A. Azimov, director of the Institute of Zoology, for continued collaboration and assistance. M. J. Ryan and an anonymous reviewer provided helpful suggestions on a draft of the manuscript.
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Sues, HD., Averianov, A. Turanoceratops tardabilis—the first ceratopsid dinosaur from Asia. Naturwissenschaften 96, 645–652 (2009). https://doi.org/10.1007/s00114-009-0518-9
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DOI: https://doi.org/10.1007/s00114-009-0518-9